Summary
of status, habitat needs, concerns, objectives, and recommended action
items, derived from species account authors and 6/23/00 meeting.
. SUBSPECIES
STATUS: None
presently observed by AOU. Grinnell
and Miller (1944) recognized Certhia familiaris zelotes and Certhia
familiaris occidentalis in California.
. MANAGEMENT
STATUS:No
official status
. RANGE
MAPS (California):
. Historical
distribution maps (subspecies): Grinnell and Miller
. BBS
map
. Christmas
bird count map
. I.Historical
references:
. Grinnell
and Miller (1944) distinguished two Brown Creeper subspecies in California.Certhia
familiaris zelotes, is a Transition Zone resident, with “partial and
irregular emigration in autumn” to lower elevations.It
was considered common “under optimum habitat conditions,” but they state
that “removal of old forest likely has reduced aggregate population.”Its
range covered almost the entire length of the state:east
of the coast belt and west of the southeastern deserts.In
the north, it extended east to the Warner Mountains, Modoc County; west
to include Siskiyou and Salmon mountains, Siskiyou County.It
extended south along the inner Coast Ranges, nearly to the Strait of Carquinez.Its
range included the entire Sierra Nevada continuously, with patchy distribution
south of Tehachapi, on higher mountains as far as Cuyamaca Mountains, San
Diego County.Grinnell and Miller
(1944) described its habitat as mature forest, particularly conifers (incense
cedar, white, red and douglas firs, yellow, jeffrey and lodgepole pines),
but also deciduous trees (black oak, aspen, cottonwood, alder; in winter:valley,
live and blue oaks).
. The
range forCerthia familiaris occidentaliswas
described as the coastal strip south of the Oregon line in Del Norte County
and as far south as Moneterey County, from sea level up to 3700 feet.It
was found in the “densest and oldest forests available,” particularly original
stands of coast redwood (Grinnell and Miller 1944).
. II.Current
breeding distribution:
. The
Brown Creeper is a “fairly common resident within the forests of its breeding
range” (Small 1994).It breeds south
of the Oregon border through the Klamath Mountains and the Northern Coast
Range to San Francisco Bay; from San Francisco Bay south, in the Southern
Coast Range to Morro Bay, and in the San Rafael Mountains (Small 1994).It
also breeds in the mountains of the northern Modoc Plateau, the Warner
Moutnains and the White Mountainns, the Inyo Mountains, the southern Cascades
south to northern Kern county and the Mt. Pinos area mountains.In
southern California it is found in the San Gabriel, San Bernardino, San
Jacinto and Santa Rosa mountains and at upper elevations in San Diego County
(Small 1994).
. BBS
relative abundance data (1966 to 1996) by region in California:
. AreaRelative
abundance
. Sierra
Nevada4.3
. South
Pacific Rainforests3.9
. Los
Angeles Ranges1.2
. Pitt-Klamath
Plateau1.1
. California
Foothills0.6
. Basin
and Range0.06
. ECOLOGY:
. I.Average
territory size:In a white cedar
bog in Michigan, territory size ranged from 2.3 to 6.4 hectares (Davis
1978).
. II.Time
of occurrence and seasonal movements.
. A.Arrival
date on breeding grounds:
. The
breeding season in California, based on nest records, ranges from April
16 through July 11; Bent (1964) notes that the height of the breeding season
as May 19 through June 11 (17 nest records found).
B.Departure
from breeding grounds:
. Some
Brown Creepers at higher elevations move downslope in the fall, with great
variation in numbers migrating from year to year (Small 1994).In
some years, during “sporadic fall migrations,” Brown Creepers disperse
to “the deserts, the valley floors and the coast” (Small 1994).Fall
dispersal may occur as early as late August, with maxium numbers recorded
during October (Small 1994).
. C.Spring
migration period:Information on
spring movement is not available (Small 1994).
.
D.Fall
migration period.See B.
. E.Extent
of wintering in California:from
Christmas bird count 1999
. Countynumber
reportedNumber/party
hour
. Big
Sur450.508
. Santa
Cruz970.433
. Woodfords90.250
. Ano
Nuevo310.248
. Yosemite
Natl Pk110.227
. Mineral
CA40.211
. Tehachapi80.190
. Oakland530.168
. Crystal
Springs270.167
. Sierra
Valley50.165
. San
Francisco220.146
. Santa
Rosa180.143
. Arcata280.138
. Moss
Landing350.118
. Grass
Valley160.113
. Monterey
Peninsula200.109
. Palo
Alto250.106
. Clear
Lake80.101
.
.
. III.Migration
stop-over needs/characteristics:
. A.Stop-over
period:No information.
. B.Habitat
use:No information.
. C.Routes:No
information.
. IV.Nest
type:
. Nests
are built behind a loose slab of bark still attached to a living or dead
tree; the nest is a hammock-like cup with a foundation of twigs, leaves,
bark shreds; and lined with finer bark, grasses, feathers, mosses.The
female builds the nest (Franzreb 1985).Building
may take a month (Franzreb 1985).Brown
Creepers may occasionally nest in holes in trees if loose bark is not available
(Harrison 1979).
. V.Foraging
strategy:
. Brown
Creepers use their slightly decurved bill to pick food items from cracks
and crevices and off the bark surface (Davis 1978).Typically,
Brown Creepers climb upward along the trunk, often spiraling around the
trunk, then drop down to the base of another tree when they reach a certain
height (Bent 1964).Birds have been
observed starting 1 meter off the ground, working their way up the trunk
to within 1-3 meters of the tree top, then flying to the trunk of another
tree (Franzreb 1985).The point at
which birds switched to another tree often coincided with an increase in
branch density, probably making it difficult for the bird to maneuver on
the trunk (Franzreb 1985).Peck-probe
was the most common foraging strategy (92.5% of observations), followed
by gleaning (6.9% of observations) and hawking (0.6% of observations) (Franzreb
1985).
. Brown
Creepers forage predominantly on the bole (98%) of a tree, spending more
time foraging in the lower bole (67%) than the upper bole (19%) (Weikel
et al. 1999).Creepers are sometimes
observed in the lower crown but not in the upper crown (Weikel et al. 1999).
. VI.Displays:
. In
courtship displays a male may “launch out from a tree and at top speed
twine around another tree, or weave in and out among the surrounding tree
and branches” (Bent 1964).
. Courtship
feeding of the female by the male takes place until egg hatching (Davis
1978).
. In
Michigan, territorial singing occurred commonly from April until the young
fledged (Davis 1978).
. VII.Social
Organization:
. A.Typical
breeding densities:In
western Oregon, Brown Creeper total density during the breeding season
increased with douglas-fir stand age:0
per 40 hectares in 10 year-old stands; 3 per 40 hectares in 35 year-old
stands; 8 per 40 hectares in 75 year-old stands; 19 per 40 hectares in
110 year-old stands; 46 per 40 hectares in 200 year-old stands (Mannan
et al. 1980).In old-growth hemlock-white
pine-hardwood forest (Appalachian Plateau, Pennsylvania) there were 1.58
breeding territories per 10 hectares (Haney 1999).
. B.Mating
system: Monogamous (Davis 1978)
. C.Delayed
breeding: No information.
. D.Post
fledging biology of offspring:Fledgling
groups initially remain within a 500-meter radius of the former nest (Davis
1978).At seventeen days after fledging,
young creepers still occasionally begged and were fed (Davis 1978).Bent
(1964) reports that young of the year may be attended by adults up to the
first week in September.However,
Davis did not find any family groups after mid-July (1978).
. E..Post
breeding social behavior:In winter
Brown Creepers are found with flocks of feeding bush-tits, kinglets, chickadees
and nuthatches (Bent 1964).
. VIII.Clutch
size:From 4 to 8 eggs to a clutch,
most commonly 5 or 6 (Bent 1964)
. IX.Incubating
sex: Both sexes.
. X.Incubation
period:15 days (Davis 1978). Incubation
begins after the clutch is complete (Davis 1978).
. XI.Nestling
period: Bent (1964) reports that young leave the nest within 13 - 14 days
of hatching.Davis (1978) reports
that they leave the nest 15-16 days after hatching.
. XII.Development
at hatching: Altricial (Davis 1978).
. XIII.Number of broods: